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In May 1944, five years after the declaration of [[World War II]], three Royal Homing Pigeon Service [[war pigeon]]s are flying across the [[English Channel]] with the [[White Cliffs of Dover]] in sight, carrying vital messages to [[Great Britain]]. Despite the poor weather conditions the pigeons have nearly reached their destination. They are, however, suddenly ambushed and attacked by a [[Axis forces]] enemy [[peregrine falcon]] named General Von Talon; two of the pigeons are instantly killed, yet the third, Mercury, is taken as a [[prisoner of war]].
Elsewhere, a small [[Common Wood Pigeon|wood pigeon]] named Valiant is watching an [[Allies of World War II|Allied forces]] propaganda film in his local bar (an overturned rowing boat) in West Nestington. He is best friends with Felix, an old [[seagull]] with a peg leg and the local barman. Wing Commander Gutsy, a [[war hero]], flies into the bar, informing everyone that signups are scheduled the next day in [[Trafalgar Square]], London. Valiant flies off to London, bidding his mother and Felix goodbye. In London, Valiant meets an unhygienic [[slacker]] pigeon named Bugsy, who is being hunted by two [[magpie]] thugs, after having tricked them at a [[shell game]]. In order to escape the thugs, he signs up with Valiant.
The recruits, Valiant, Bugsy, Lofty, an intellectual red pigeon, and Toughwood and Tailfeather, two muscular but dim-witted twin brothers, form Royal Homing Pigeon Service Squad F, and are sent to a [[recruit training]] facility. Under the command of Sergeant Monty, who declares that he will toughen them up for the RHPS, the training begins. Meanwhile, Von Talon and his henchmen, Cufflingk and Underlingk, try numerous attempts to discover the message's departure location. However, Mercury refuses to tell, despite the tortures inflicted upon him, such as irritating him with yodeling music and injecting him with [[truth serum]], before Mercury accidentally reveals the location: Saint-Pierre.
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