Jump to content

Chenopodium: Difference between revisions

From Wikipedia, the free encyclopedia
Browse history interactively
← Previous edit
Content deleted Content added
VisualWikitext
Eubot (talk | contribs)
259,670 edits
m Removed taxobox colour. See User:Eubot/Removing colours from taxoboxen.
m Updated short description
Tags: Mobile edit Mobile app edit iOS app edit App description change
 
(186 intermediate revisions by more than 100 users not shown)
Line 1: Line 1:
{{Short description|Genus of flowering plants in the amaranth family}}
:''"Goosefoot" redirects here. The unrelated [[Smearwort]] (''Aristolochia rotunda'') is sometimes called "Mercury Goosefoot".
{{Taxobox
{{hatgrp|
{{redirect|Goosefoot}}
| name = Goosefoots, genus ''Chenopodium''
{{redirect|Rhagodia|the arachnid genus|Rhagodia (arachnid){{!}}''Rhagodia'' (arachnid)}}
| image = Melganzenvoet bloeiwijze Chenopodium album.jpg
| image_width = 250px
| image_caption = Fat Hen (''Chenopodium album'')
| regnum = [[Plant]]ae
| divisio = [[Flowering plant|Magnoliophyta]]
| classis = [[Magnoliopsida]]
| ordo = [[Caryophyllales]]
| familia = [[Amaranthaceae]]
| genus = '''''Chenopodium'''''
| subdivision_ranks = species
| subdivision = See text
}}
}}
{{More citations needed|date=March 2021}}
{{Automatic taxobox
|image = Chenopodium berlandieri NPS-1.jpg
|image_caption = ''[[Chenopodium berlandieri]]''
|display_parents = 2
|taxon = Chenopodium
|authority = [[Carl Linnaeus|L.]] (1753)
|subdivision_ranks = [[Species]]
|subdivision = 132; see [[#Selected species|text]]
|subdivision_ref = <ref name = powo>[https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:60437273-2 ''Chenopodium'' L.] ''[[Plants of the World Online]]''. Retrieved 4 April 2024.</ref>
|synonyms =
*''Agatophyton'' {{small|Fourr. (1869)}}
*''Blitum'' {{small|Hill (1757), nom. illeg.}}
*''Botrys'' {{small|Rchb. ex Nieuwl. (1914)}}
* ''Einadia'' <small>Raf. (1838)</small>
* ''Rhagodia'' <small>R.Br. (1810)</small>
* ''Vulvaria'' <small>Bubani (1897), nom. illeg.</small>
* ''Chenopodium'' sect. ''Leprophyllum'' <small>Dumort.</small>
* ''Chenopodium'' sect. ''Chenopodiastrum'' <small>Moq.</small>
|synonyms_ref = <ref name = powo/><ref name="Fuentes"/>
}}

'''''Chenopodium''''' is a [[genus]] of numerous [[species]] of [[perennial]] or [[annual plant|annual]] [[herbaceous]] [[flowering plant]]s known as the '''goosefoot''', which occur almost anywhere in the world.<ref name="Fl China" /> It is placed in the [[family (biology)|family]] [[Amaranthaceae]] in the [[APG II system]]; older classification systems, notably the widely used [[Cronquist system]], separate it and its relatives as [[Chenopodiaceae]],<ref>{{cite EB1911|wstitle=Chenopodium|volume=6|page=80}}</ref> but this leaves the rest of the Amaranthaceae [[polyphyletic]]. However, among the Amaranthaceae, the genus ''Chenopodium'' is the namesake member of the [[subfamily]] [[Chenopodioideae]].<ref name="Fuentes" />

==Description==
[[File:Chenopodium album (4032134406).jpg|thumb|left|upright=.8|White goosefoot (''[[Chenopodium album]]'')]]

The species of ''Chenopodium'' (s.str., description according to Fuentes et al. 2012)<ref name="Fuentes" /> are [[Annual plant|annual]] or [[perennial]] [[herbs]], [[shrubs]] or small [[trees]].<ref name=":1">{{Cite book|last=Taylor|first=Ronald J.|url=https://www.worldcat.org/oclc/25708726|title=Sagebrush Country: A Wildflower Sanctuary|publisher=Mountain Press Pub. Co|year=1994|isbn=0-87842-280-3|edition=rev.|location=Missoula, MT|pages=64|language=en|oclc=25708726|orig-year=1992}}</ref> They generally rely on [[alkaline soil]].<ref name=":1" /> They are nonaromatic, but sometimes fetid. The young stems and leaves are often densely covered by vesicular globose hairs, thus looking [[Epicuticular wax#Farina|farinose]]. Characteristically, these [[trichomes]] persist, collapsing later and becoming cup-shaped.
The branched [[Plant stem|stems]] grow erect, ascending, prostrate or scrambling. Lateral branches are alternate (the lowermost ones can be nearly opposite). The alternate or opposite [[leaves]] are petiolate. Their thin or slightly fleshy leaf blade is linear, rhombic or triangular-hastate, with entire or dentate or lobed margins.<ref name="Fuentes" />

[[Inflorescence]]s are standing terminal and lateral. They consist of spicately or paniculately arranged glomerules of flowers. Plants are [[monoecious]] (rarely [[dioecious]]). In monoecious plants flowers are dimorphic or [[pistillate]]. Flowers consist of (4–) 5 [[perianth]] segments connate, basally or close to the middle, usually membranous margined and with a roundish to keeled back; almost always 5 stamens, and one ovary with 2 stigmas.<ref name="Fuentes" />

In fruit, perianth segments become sometimes coloured, but mostly keep unchanged, somewhat closing over or spreading from the fruit. The [[pericarp]] is membranous or sometimes succulent, adherent to or loosely covering the seed. The horizontally oriented seeds are depressed-globular to lenticular, with rounded to subacute margin. The black seed coat is almost smooth to finely striate, rugulose or pitted.<ref name="Fuentes" />

== Ecology ==
Certain species grow in large [[thicket]]s, providing cover for small animals. Goosefoot foliage is used as food by the [[caterpillar]]s of certain [[Lepidoptera]].{{Citation needed|date=August 2020}} The seeds are eaten by many [[bird]]s, such as the [[yellowhammer]] (''Emberiza citrinella'') of Europe or the [[white-winged fairy-wren]] (''Malurus leucopterus'') of [[Australia]].{{Citation needed|date=August 2020}} Goosefoot [[plant pathogen|pathogens]] include the [[positive-sense ssRNA virus]]es{{snd}}[[apple stem grooving virus]], [[sowbane mosaic virus]] and [[tobacco necrosis virus]].{{Citation needed|date=August 2020}}

==Uses and human importance==
[[File:Quinoa cuit.JPG|thumb|Cooked [[quinoa]] (''C. quinoa'') seeds]]

The genus ''Chenopodium'' contains several plants of minor to moderate importance as food crops as [[leaf vegetable]]s &ndash; used like the closely related [[spinach]] (''Spinacia oleracea'') and similar plants called ''quelite'' in [[Mexico]]{{snd}}and [[pseudocereal]]s.{{Citation needed|date=August 2020}} These include [[white goosefoot]] (''C.&nbsp;album''), ''[[kañiwa]]'' (''C.&nbsp;pallidicaule'') and [[quinoa]] (''C.&nbsp;quinoa''). On the [[Greece|Greek]] island of [[Crete]], tender shoots and leaves of a species called ''krouvida'' ({{langr|el|κρουβίδα}}) or ''psarovlito'' ({{langr|el|ψαρόβλητο}}) are eaten by the locals, boiled or steamed.{{Citation needed|date=August 2020}} As studied by [[Bruce D. Smith]], [[Kristen Gremillion]] and others, goosefoots have a history of culinary use dating back to 4000 BC or earlier, when [[pitseed goosefoot]] (''C.&nbsp;berlandieri'') was a staple crop in the Native American [[Eastern Agricultural Complex]],{{Citation needed|date=August 2020}} and when white goosefoot was apparently used by the [[Ertebølle culture]] of [[Europe]].{{Citation needed|date=August 2020}} Members of the eastern European [[Yamna culture|Yamnaya culture]] also harvested white goosefoot as an apparent cereal substitute to round out an otherwise mostly meat and dairy diet c.''&nbsp;''3500–2500''&nbsp;''BC.<ref>{{Cite book|last=Anthony|first=David|title=The horse, the wheel, and language|publisher=Princeton university press|year=2007|pages=302–303}}</ref>

There is increased interest in particular in goosefoot seeds today, which are suitable as part of a [[gluten-free diet]].{{Citation needed|date=August 2020}} [[Quinoa oil]], extracted from the seeds of ''C. quinoa'', has similar properties, but is superior in quality, to [[corn oil]].{{Citation needed|date=August 2020}} Oil of chenopodium is extracted from the seeds of [[epazote]], which is not in this genus anymore.{{Citation needed|date=August 2020}} [[Shagreen]] leather was produced in the past using the small, hard goosefoot seeds.{{Citation needed|date=August 2020}} ''C.&nbsp;album'' was one of the main [[model organism]]s for the [[molecular biological]] study of [[chlorophyllase]].{{Citation needed|date=August 2020}}

Goosefoot [[pollen]], in particular of the widespread and usually abundant ''C.&nbsp;album'', is an [[allergen]] to many people and a common cause of [[hay fever]].<ref name=":0">{{Cite web|date=2017-10-13|title=Amaranthus and Chenopodium|url=https://alabamaallergy.com/2017/10/13/amaranthus-and-chenopodium/|access-date=2021-03-03|website=alabamaallergy.com|language=en-US}}</ref> The same species, as well as some others, have seeds which are able to persist for years in the [[soil seed bank]].{{Citation needed|date=August 2020}} Many goosefoot species are thus significant [[weed]]s, and some have become [[invasive species]].<ref name=":0" />

In Australia, the larger ''Chenopodium'' species are among the plants called "bluebushes".{{Citation needed|date=August 2020}} According to the 1889 book ''The Useful Native Plants of Australia'', ''Chenopodium auricomum'' "is another of the salt-bushes, which, besides being invaluable food for stock, can be eaten by man. All plants of the Natural Order Chenopodiaceae (Salsolacese) are more or less useful in this respect." The book goes on to give the following account from the ''Journal de la Ferme et des Maisons de campagne'':<ref>{{cite book | author=J. H. Maiden | year=1889 | title=The useful native plants of Australia : Including Tasmania | publisher= Turner and Henderson, Sydney | pages=15–16| url=https://archive.org/details/usefulnativeplan00maid/page/15}}</ref>

<blockquote>We have recently gathered an abundant harvest of leaves from two or three plants growing in our garden. These leaves were put into boiling water to blanch them, and they were then cooked as an ordinary dish of spinach, with this difference in favour of the new plant, that there was no occasion to take away the threads which are so disagreeable in chicory, sorrel, and ordinary spinach. We partook of this dish with relish—the flavour—analogous to spinach, had something in it more refined, less grassy in taste. The cultivation is easy: sow the seed in April (October) in a well-manured bed, for the plant is greedy; water it. The leaves may be gathered from the time the plant attains 50 centimetres (say 20 inches) in height. They grow up again quickly. In less than eight days afterwards another gathering may take place, and so on to the end of the year.</blockquote>

==Systematics==
The genus ''Chenopodium'' was described by [[Carl Linnaeus]] in 1753 (In: ''Species Plantarum'', Vol. 1, p.&nbsp;218–222). Type species is ''[[Chenopodium album]]''. This generic name is derived from the particular shape of the leaf, which is similar to a goose's foot: from [[Greek language|Greek]] χήν (''chen''), "goose" and πούς (''pous''), "foot" or {{langr|el|ποδίον}} (''podion''), "little foot".

In its traditional circumscription, ''Chenopodium'' comprised about 170 species.<ref name="Fl China" /> Phylogenetic research revealed, that the genus was highly [[polyphyletic]] and did not reflect how species were naturally related. Therefore, a new classification was necessary. Mosyakin & Clemants (2002, 2008) separated the glandular species as genus ''[[Dysphania (plant)|Dysphania]]'' (which includes [[epazote]]) and ''[[Teloxys]]'' in tribe [[Dysphanieae]]. Fuentes-Bazan et al. (2012) separated many species to genera ''[[Blitum]]'' (in tribe [[Anserineae]]), ''[[Chenopodiastrum]]'', ''[[Lipandra]]'', and ''[[Oxybasis]]'' (like ''Chenopodium'' in tribe [[Atripliceae]]). They included ''Rhagodia'' and ''Einadia'' in ''Chenopodium''.<ref name="Fuentes" />


===Species===
'''''Chenopodium''''' is a genus of about 150 species of [[flowering plant]]s, known generically as the '''Goosefoots'''. It contains several plants of minor to moderate importance as [[food]] crops, both [[leaf vegetable]]s and pseudo-[[cereal]]s, including [[Quinoa]], [[Kañiwa]], [[Fat Hen]], [[Good King Henry]], and [[Epazote]]. Goosefoots are used as food plants by the [[larva]]e of some [[Lepidoptera]] species; see [[list of Lepidoptera that feed on Chenopodium]].
132 species are currently accepted.<ref name = powo/>
[[File:Starr 010206-0253 Chenopodium oahuense.jpg|thumb|{{okina}}Āheahea (''[[Chenopodium oahuense]]'')]]
[[File:Rhagodia candolleana.jpg|thumb|''[[Chenopodium candolleanum]]'']]
[[File:좀명아주.JPG|thumb|''[[Chenopodium ficifolium]]'']]
[[File:MagentaSpreen Chenopodium giganteum EdibleOffice.jpg|thumb|''[[Chenopodium giganteum]]'']]
{{div col|colwidth=35em}}
* ''[[Chenopodium acerifolium]]'' {{small|Andrz.}}
* ''[[Chenopodium aciculare]]'' {{small|(Paul G.Wilson) S.Fuentes & Borsch}}
* ''[[Chenopodium acuminatum]]'' <small>Willd.</small><!-- BiolConserv137:248. -->
* ''[[Chenopodium adpressifolium]]'' {{small|Pandeya & A.Pandeya}}
* ''[[Chenopodium albescens]]'' {{small|Small}}
* ''[[Chenopodium album]]'' {{small|L.}} &ndash; white goosefoot, nickel greens, dungweed, ''bathua'', ''chandali'', ''chandaliya'', fat hen, lamb's quarters, pigweed
** ''Chenopodium album'' var. ''album'' (synonym ''[[Chenopodium strictum]]'' <small>Roth</small>)<!-- BiolConserv137:248. -->
* ''[[Chenopodium allanii]]'' {{small|Aellen}}
* ''[[Chenopodium atripliciforme]]'' {{small|Murr}}
* ''[[Chenopodium atrovirens]]'' {{small|Rydb.}} &ndash; dark goosefoot, pinyon goosefoot
* ''[[Chenopodium attenuatum]]'' {{small|Charit.}}
* ''[[Chenopodium aureum]]'' {{small|Benet-Pierce}} &ndash; golden goosefoot
* ''[[Chenopodium auricomiforme]]'' {{small|Murr & Thell.}}
* ''[[Chenopodium auricomum]]'' {{small|Lindl.}} &ndash; Queensland bluebush
* ''[[Chenopodium ayare]]'' {{small|Toro Torr.}}
* ''[[Chenopodium baccatum]]'' {{small|Labill.}} (Syn. ''Rhagodia baccata'')
* ''[[Chenopodium benthamii]]'' {{small|Iamonico & Mosyakin}} (Syn.: ''Rhagodia latifolia'')
* ''[[Chenopodium berlandieri]]'' {{small|Moq.}} &ndash; pitseed goosefoot, southern huauzontle, lambsquarters
** ''Chenopodium berlandieri'' var. ''berlandieri'' (synonym ''[[Chenopodium nuttalliae]]'' {{small|Saff.}}) &ndash; huauzontle, chia roja, quelit
* ''[[Chenopodium betaceum]]'' {{small|Andrz.}}
* ''[[Chenopodium × binzianum]]'' {{small|Aellen & Thell.}}
* ''[[Chenopodium × bohemicum]]'' {{small|F.Dvořák}}
* ''[[Chenopodium × borbasii]]'' {{small|Murr}}
* ''[[Chenopodium brandegeeae]]'' {{small|Benet-Pierce}} &ndash; Brandegee's goosefoot
* ''[[Chenopodium bryoniifolium]]'' <small>Bunge</small> – Korean goosefoot<ref>{{Cite book|url=http://www.forest.go.kr/kna/special/download/English_Names_for_Korean_Native_Plants.pdf|title=English Names for Korean Native Plants|publisher=[[Korea National Arboretum]]|year=2015|isbn=978-89-97450-98-5|location=Pocheon|pages=407|access-date=24 December 2016|via=[[Korea Forest Service]]|url-status=dead|archive-url=https://web.archive.org/web/20170525105020/http://www.forest.go.kr/kna/special/download/English_Names_for_Korean_Native_Plants.pdf|archive-date=25 May 2017}}</ref>
* ''[[Chenopodium candolleanum]]'' {{small|(Moq.) S.Fuentes & Borsch}} (Syn.: ''Rhagodia candolleana'')
* ''[[Chenopodium carnosulum]]'' {{small|Moq.}}
* ''[[Chenopodium chaldoranicum]]'' {{small|Rahimin. & Ghaemm.}}
* ''[[Chenopodium cordobense]]'' {{small|Aellen}}
* ''[[Chenopodium cornutum]]'' {{small|(Torr.) Benth. & Hook.f. ex S.Watson}}
* ''[[Chenopodium × covillei]]'' {{small|Aellen}}
* ''[[Chenopodium crusoeanum]]'' {{small|Skottsb.}}
* ''[[Chenopodium cuneifolium]]'' {{small|Vahl}}
* ''[[Chenopodium curvispicatum]]'' {{small|Paul G.Wilson}}
* ''[[Chenopodium cyanifolium]]'' {{small|Pandeya, Singhal & A.K.Bhatn.}}
* ''[[Chenopodium cycloides]]'' {{small|A.Nelson}} &ndash; sandhill goosefoot
* ''[[Chenopodium × dadakovae]]'' {{small|F.Dvořák}}
* ''[[Chenopodium desertorum]]'' {{small|(J.M.Black) J.M.Black}} &ndash; desert goosefoot
* ''[[Chenopodium desiccatum]]'' {{small|A.Nelson}} &ndash; narrowleaf goosefoot
* ''[[Chenopodium detestans]]'' {{small|Kirk}} &ndash; New Zealand fish-guts plant
* ''[[Chenopodium diversifolium]]'' {{small|(Aellen) F.Dvořák}}
* ''[[Chenopodium drummondii]]'' {{small|(Moq.) S.Fuentes & Borsch}} (Syn.: ''Rhagodia drummondii'')
* ''[[Chenopodium eastwoodiae]]'' {{small|Benet-Pierce}} &ndash; Eastwood's goosefoot
* ''[[Chenopodium eremaeum]]'' {{small|(Paul G.Wilson) S.Fuentes & Borsch}} (Syn.: ''Rhagodia eremaea'')
* ''[[Chenopodium erosum]]'' <small>R.Br.</small>
* ''[[Chenopodium eustriatum]]'' {{small|F.Dvořák}}
* ''[[Chenopodium × fallax]]'' {{small|(Aellen) F.Dvořák}}
* ''[[Chenopodium ficifoliiforme]]'' {{small|F.Dvořák}}
* ''[[Chenopodium ficifolium]]'' {{small|Sm.}} &ndash; fig-leaved goosefoot, small goosefoot
* ''[[Chenopodium flabellifolium]]'' {{small|Standl.}} &ndash; San Martin Island goosefoot, flabelliform goosefoot
* ''[[Chenopodium foggii]]'' {{small|Wahl}} &ndash; Fogg's goosefoot
* ''[[Chenopodium fremontii]]'' {{small|S.Watson}} &ndash; Fremont's goosefoot
* ''[[Chenopodium frutescens]]'' {{small|C.A.Mey.}}
* ''[[Chenopodium × fursajevii]]'' {{small|Aellen & Iljin}}
* ''[[Chenopodium gaudichaudianum]]'' {{small|(Moq.) Paul G.Wilson}}
* ''[[Chenopodium giganteum]]'' <small>D.Don</small> (synonym ''[[Chenopodium formosanum]]'' {{small|Koidz.}}) &ndash; tree spinach, red quinoa, djulis<!-- BiolConserv137:248. -->
* ''[[Chenopodium griseochlorinum]]'' {{small|F.Dvořák}}
* ''[[Chenopodium grubovii]]'' {{small|Lomon. & Uotila}}
* ''[[Chenopodium × gruellii]]'' {{small|Aellen}}
* ''[[Chenopodium harae]]'' {{small|Sukhor.}}
* ''[[Chenopodium hastatifolium]]'' {{small|Pandeya & A.Pandeya}}
* ''[[Chenopodium × haywardiae]]'' {{small|Murr}}
* ''[[Chenopodium hederiforme]]'' {{small|(Murr) Aellen}}
* ''[[Chenopodium hians]]'' {{small|Standl.}}
* ''[[Chenopodium hircinum]]'' {{small|Schrad.}}
* ''[[Chenopodium hoggarense]]'' {{small|Uotila & C.Chatel.}}
* ''[[Chenopodium howellii]]'' {{small|Benet-Pierce}} &ndash; Howell's goosefoot
* ''[[Chenopodium hubbardii]]'' {{small|Aellen}}
* ''[[Chenopodium × humiliforme]]'' {{small|(Murr) F.Dvořák}}
* ''[[Chenopodium iljinii]]'' {{small|Golosk.}}
* ''[[Chenopodium incanum]]'' {{small|(S.Watson) A.Heller}} &ndash; mealy goosefoot
* ''[[Chenopodium incognitum]]'' {{small|Wahl}}
* ''[[Chenopodium indicum]]'' {{small|T.K.Paul}}
* ''[[Chenopodium iranicum]]'' {{small|(Aellen) Hamdi & Malekloo}}
* ''[[Chenopodium × jedlickae]]'' {{small|F.Dvořák}}
* ''[[Chenopodium × jehlikii]]'' {{small|F.Dvořák}}
* ''[[Chenopodium karoi]]'' {{small|(Murr) Aellen}}
* ''[[Chenopodium khorasanica]]'' {{small|Hamdi & Malekloo}}
* ''[[Chenopodium lenticulare]]'' {{small|Aellen}}
* ''[[Chenopodium × leptophylliforme]]'' {{small|Aellen}}
* ''[[Chenopodium leptophyllum]]'' {{small|(Moq.) Nutt. ex S.Watson}} &ndash; narrowleaf goosefoot
* ''[[Chenopodium × linciense]]'' {{small|Murr}}
* ''[[Chenopodium lineatum]]'' {{small|Benet-Pierce}} &ndash; Mono goosefoot
* ''[[Chenopodium littoreum]]'' {{small|Benet-Pierce & M.G.Simpson}} &ndash; coastal goosefoot
* ''[[Chenopodium lobodontum]]'' {{small|H.Scholz}}
* ''[[Chenopodium loureiroi]]'' {{small|Steud.}}
* ''[[Chenopodium luteorubrum]]'' {{small|Mandák & Lomon.}}
* ''[[Chenopodium luteum]]'' {{small|Benet-Pierce}} &ndash; yellow goosefoot
* ''[[Chenopodium × mendelii]]'' {{small|F.Dvořák}}
* ''[[Chenopodium moquinianum]]'' {{small|Aellen}}
* ''[[Chenopodium mucronatum]]'' {{small|Thunb.}}
* ''[[Chenopodium neomexicanum]]'' {{small|Standl.}} &ndash; New Mexico goosefoot
* ''[[Chenopodium nesodendron]]'' {{small|Skottsb.}}
* ''[[Chenopodium nevadense]]'' {{small|Standl.}} &ndash; Nevada goosefoot
* ''[[Chenopodium nitens]]'' {{small|Benet-Pierce & M.G.Simpson}}
* ''[[Chenopodium nitrariaceum]]'' <small>(F.Muell.) F.Muell. ex Benth.</small> &ndash; nitre goosefoot
* ''[[Chenopodium nitens]]'' {{small|Benet-Pierce & M.G.Simpson}} &ndash; shiny goosefoot
* ''[[Chenopodium novopokrovskyanum]]'' {{small|(Aellen) Uotila}}
* ''[[Chenopodium nutans]]'' {{small|(R.Br.) S.Fuentes & Borsch}} (Syn.: ''Rhagodia nutans'')
* ''[[Chenopodium oahuense]]'' {{small|(Meyen) Aellen}} &ndash; ''{{okina}}Āheahea'' ([[Hawaii|Hawai{{okina}}i]])
* ''[[Chenopodium obscurum]]'' {{small|Aellen}}
* ''[[Chenopodium opulifolium]]'' <small>Schrad. ex W.D.J.Koch & Ziz</small> &ndash; grey goosefoot<!-- BiolConserv137:248. -->
* ''[[Chenopodium pallescens]]'' {{small|Standl.}} &ndash; pallid goosefoot
* ''[[Chenopodium pallidicaule]]'' {{small|Aellen}} &ndash; ''kañiwa'', "cañahua"
* ''[[Chenopodium palmeri]]'' {{small|Standl.}} &ndash; Palmer's goosefoot
* ''[[Chenopodium pamiricum]]'' {{small|Iljin}}
* ''[[Chenopodium parabolicum]]'' {{small|(R.Br.) S.Fuentes & Borsch}} (Syn.: ''Rhagodia parabolica'')
* ''[[Chenopodium × paradoxum]]'' {{small|Mandák}}
* ''[[Chenopodium parryi]]'' {{small|Standl.}} &ndash; Parry's goosefoot
* ''[[Chenopodium perttii]]'' {{small|Sukhor.}}
* ''[[Chenopodium petiolare]]'' {{small|Kunth}}
* ''[[Chenopodium philippianum]]'' {{small|Aellen}}
* ''[[Chenopodium phillipsianum]]'' {{small|Aellen}}
* ''[[Chenopodium pilcomayense]]'' {{small|Aellen}}
* ''[[Chenopodium × podperae]]'' {{small|F.Dvořák}}
* ''[[Chenopodium polygonoides]]'' {{small|(Murr) Aellen}}
* ''[[Chenopodium × praeacutum]]'' {{small|Murr}}
* ''[[Chenopodium pratericola]]'' <small>Rydb.</small> &ndash; pale goosefoot, desert goosefoot, narrowleaf goosefoot<!-- BiolConserv137:248. -->
* ''[[Chenopodium preissii]]'' {{small|(Moq.) Diels}} (Syn. ''Rhagodia preissii'')
* ''[[Chenopodium × preissmannii]]'' {{small|Murr}}
* ''[[Chenopodium × pseudoleptophyllum]]'' {{small|Aellen}}
* ''[[Chenopodium × pseudostriatum]]'' {{small|(Zschacke) Druce}}
* ''[[Chenopodium pueblense]]'' {{small|H.S.Reed}}
* ''[[Chenopodium quinoa]]'' {{small|Willd.}} &ndash; quinoa
* ''[[Chenopodium × reynieri]]'' {{small|Ludw. & Aellen}}
* ''[[Chenopodium robertianum]]'' {{small|Iamonico & Mosyakin}} (Syn.: ''Rhagodia hastata'')
* ''[[Chenopodium ruiz-lealii]]'' {{small|Aellen}}
* ''[[Chenopodium sanctae-clarae]]'' {{small|Johow}}
* ''[[Chenopodium sancti-ambrosii]]'' {{small|Skottsb.}}
* ''[[Chenopodium sandersii]]'' {{small|Benet-Pierce}} &ndash; Sander's goosefoot
* ''[[Chenopodium santoshei]]'' {{small|Pandeya, Singhal & A.K.Bhatn.}}
* ''[[Chenopodium scabricaule]]'' {{small|Speg.}}
* ''[[Chenopodium simpsonii]]'' {{small|Benet-Pierce}} &ndash; Simpson's goosefoot
* ''[[Chenopodium × smardae]]'' {{small|F.Dvořák}}
* ''[[Chenopodium sonorense]]'' {{small|Benet-Pierce & M.G.Simpson}} &ndash; Sonoran goosefoot
* ''[[Chenopodium sosnowskyi]]'' {{small|Kapeller}}
* ''[[Chenopodium spegazzinii]]'' {{small|F.Dvořák}}
* ''[[Chenopodium spinescens]]'' {{small|(R.Br.) S.Fuentes & Borsch}} (Syn. ''Rhagodia spinescens'')
* ''[[Chenopodium standleyanum]]'' {{small|Aellen}} &ndash; Standley's goosefoot
* ''[[Chenopodium stenophyllum]]'' {{small|(Makino) Koidz.}}
* ''[[Chenopodium striatiforme]]'' {{small|Murr}}
* ''[[Chenopodium subficifolium]]'' {{small|(Murr) Druce}}
* ''[[Chenopodium subglabrum]]'' {{small|(S.Watson) A.Nelson}} &ndash; smooth arid goosefoot, smooth goosefoot
* ''[[Chenopodium suecicum]]'' {{small|Murr}} &ndash; green goosefoot
* ''[[Chenopodium × thellungii]]'' {{small|Murr}}
* ''[[Chenopodium tonkinense]]'' {{small|Courchet}}
* ''[[Chenopodium triandrum]]'' {{small|G.Forst.}} (Syn.: ''Rhagodia triandra'')
* ''[[Chenopodium × tridentinum]]'' {{small|Murr}}
* ''[[Chenopodium × trigonocarpum]]'' {{small|Aellen}}
* ''[[Chenopodium trigonon]]'' {{small|Schult.}} (Syn.: ''Einadia trigonos'')
* ''[[Chenopodium twisselmannii]]'' {{small|Benet-Pierce}} &ndash; Twisselmann's goosefoot, high meadow goosefoot
* ''[[Chenopodium ulbrichii]]'' {{small|Aellen}}
* ''[[Chenopodium ulicinum]]'' {{small|Gand.}}
* ''[[Chenopodium × unarii]]'' {{small|F.Dvořák}}
* ''[[Chenopodium × variabile]]'' {{small|Aellen}} (''C. album × C. berlandieri'')
* ''[[Chenopodium vulvaria]]'' {{small|L.}} &ndash; stinking goosefoot, notchweed
* ''[[Chenopodium wahlii]]'' {{small|Benet-Pierce}} &ndash; Wahl's goosefoot
* ''[[Chenopodium watsonii]]'' {{small|A.Nelson}} &ndash; Watson's goosefoot
* ''[[Chenopodium wilsonii]]'' {{small|S.Fuentes, Borsch & Uotila}} (Syn.: ''Rhagodia crassifolia'')
* ''[[Chenopodium × zahnii]]'' {{small|Murr}}
* ''[[Chenopodium zerovii]]'' {{small|Iljin}}
* ''[[Chenopodium zoellneri]]'' {{small|Aellen}}
{{div col end}}


===Excluded species===
In the [[APG II system]] (2003) it is placed in the family [[Amaranthaceae]], but earlier, for example in the [[Cronquist system]] (1981), it was placed in the family [[Chenopodiaceae]].
* ''[[Blitum]]'' (12 species):<ref name="Fuentes" />
** ''[[Blitum bonus-henricus]]'' &ndash; Good King Henry, perennial goosefoot, poor-man's asparagus, Lincolnshire spinach, markery
** ''[[Blitum californicum]]'' &ndash; California goosefoot, Indian lettuce
** ''[[Blitum capitatum]]'' &ndash; strawberry blite, blite goosefoot, strawberry goosefoot, strawberry spinach, Indian paint, Indian ink
** ''[[Blitum virgatum]]'' (Syn. ''Chenopodium foliosum'') &ndash; leafy goosefoot
* ''[[Chenopodiastrum]]'' (5 species):<ref name="Fuentes" />
** ''[[Chenopodiastrum murale]]'' &ndash; nettle-leaved goosefoot
** ''[[Chenopodiastrum simplex]]'' &ndash; giant seed goosefoot
* ''[[Dysphania (plant)|Dysphania]]'' (about 43 glandular species, as ''C. botrys, C. carinatum, C. cristatum, C. melanocarpum, C. multifidium, C. pumilio'' and more)
* ''[[Lipandra]]'' (one species):<ref name="Fuentes" />
** ''Lipandra polysperma'' &ndash; many-seeded goosefoot
* ''[[Oxybasis]]'' (5 species):<ref name="Fuentes" />
** ''[[Oxybasis chenopodioides]]'' &ndash; small red goosefoot, saltmarsh goosefoot
** ''[[Oxybasis glauca]]'' &ndash; oak-leaved goosefoot
** ''[[Oxybasis rubra]]'' &ndash; red goosefoot, coastblite goosefoot
** ''[[Oxybasis urbica]]'' &ndash; upright goosefoot
* ''[[Teloxys]]'' (one species):
** ''Teloxys aristata''
* ''[[Suaeda australis]]'' &ndash; austral seablite (as ''C. australe, C. insulare'')


==Fossil record==
In some parts of northern India, the leaves (also known as bathua or chandaliya or chandali) are used for culinary purposes
†'''''Chenopodium wetzleri''''' fossil seeds of the [[Chattian]] stage, [[Oligocene]], are known from the Oberleichtersbach Formation in the [[Rhön Mountains]], central [[Germany]].<ref>The floral change in the tertiary of the Rhön mountains (Germany) by Dieter Hans Mai - Acta Paleobotanica 47(1): 135-143, 2007.</ref>


==References==
==Selected species==
<references>
*''[[Chenopodium album]]'' (Lamb's quarters, Fat hen)
<ref name="Fuentes">Susy Fuentes-Bazan, Pertti Uotila, Thomas Borsch: ''A novel phylogeny-based generic classification for Chenopodium sensu lato, and a tribal rearrangement of Chenopodioideae (Chenopodiaceae).'' In: ''Willdenowia.'' Vol. 42, No. 1, 2012, p. 5-24.</ref>
*''[[Chenopodium ambrosoides]]'' ([[Epazote]])
<ref name="Fl China">Gelin Zhu, Sergei L. Mosyakin & Steven E. Clemants: [http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=106630 ''Chenopodium'' - In: Wu Zhengyi, Peter H. Raven, Deyuan Hong (Hrsg.): Flora of China.] Volume 5: Ulmaceae through Basellaceae. Science Press/Missouri Botanical Garden Press, Beijing/St. Louis 2003, {{ISBN|1-930723-27-X}}, p. 378-.</ref>
*''[[Chenopodium berlandieri]]'' (Pitseed Goosefoot)
</references>
*''[[Chenopodium bonus-henricus]]'' ([[Good King Henry]])
*''[[Chenopodium botrys]]'' ([[Jerusalem Oak Goosefoot]], Feathered Geranium)
*''[[Chenopodium capitatum]]'' (Strawberry-blite)
*''[[Chenopodium chenopodioides]]'' (Small Red Goosefoot)
*''[[Chenopodium ficifolium]]'' (Fig-leaved Goosefoot)
*''[[Chenopodium foliosum]]''
*''[[Chenopodium giganteum]]''
*''[[Chenopodium glaucum]]'' (Oak-leaved Goosefoot)
*''[[Chenopodium hybridum]]'' (Maple-leaved Goosefoot)
*''[[Chenopodium murale]]'' (Nettle-leaved Goosefoot)
*''[[Chenopodium opulifolium]]'' (Grey Goosefoot)
*''[[Chenopodium pallidicaule]]'' ([[Kañiwa]])
*''[[Chenopodium polyspermum]]'' (Many-seeded Goosefoot)
*''[[Chenopodium purpurascens]]'' (Purple Goosefoot)
*''[[Chenopodium quinoa]]'' ([[Quinoa]])
*''[[Chenopodium rubrum]]'' (Red Goosefoot)
*''[[Chenopodium suecicum]]'' (Green Goosefoot)
*''[[Chenopodium urbicum]]'' (Upright Goosefoot)
*''[[Chenopodium vulvaria]]'' (Stinking Goosefoot, Notch-weed)


==Further reading==
[[Category:Amaranthaceae]]
* {{cite journal |last1=Sukhorukov |first1=Alexander P. |last2=Zhang |first2=Mingli |date=2013 |title=Fruit and Seed Anatomy of ''Chenopodium'' and Related Genera (Chenopodioideae, Chenopodiaceae/Amaranthaceae): Implications for Evolution and Taxonomy |journal=PLOS ONE |volume=8 |number=4 |page=e61906 |doi=10.1371/journal.pone.0061906 |pmc=3633980 |pmid=23626750|bibcode=2013PLoSO...861906S |doi-access=free }}


{{Taxonbar|from=Q158094}}
{{Amaranthaceae-stub}}
{{Authority control}}


[[Category:Chenopodium| ]]
[[da:Gåsefod]]
[[Category:Amaranthaceae genera]]
[[de:Gänsefüße]]
[[Category:Taxa named by Carl Linnaeus]]
[[es:Chenopodium]]
[[Category:Chenopodioideae]]
[[fr:Chenopodium]]
[[he:כף האווז]]
[[hsb:Pólšica]]
[[lt:Balanda]]
[[nl:Ganzenvoet]]
[[pl:Komosa]]
[[pt:Chenopodium]]
[[ru:Марь (растение)]]

Latest revision as of 03:39, 28 December 2024

"Goosefoot" redirects here. For other uses, see Goosefoot (disambiguation). "Rhagodia" redirects here. For the arachnid genus, see Rhagodia (arachnid).

Chenopodium
Chenopodium berlandieri
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Order: Caryophyllales
Family: Amaranthaceae
Subfamily: Chenopodioideae
Tribe: Atripliceae
Genus: Chenopodium
L. (1753)
Species[1]

132; see text

Synonyms[1][2]
  • Agatophyton Fourr. (1869)
  • Blitum Hill (1757), nom. illeg.
  • Botrys Rchb. ex Nieuwl. (1914)
  • Einadia Raf. (1838)
  • Rhagodia R.Br. (1810)
  • Vulvaria Bubani (1897), nom. illeg.
  • Chenopodium sect. Leprophyllum Dumort.
  • Chenopodium sect. Chenopodiastrum Moq.

Chenopodium is a genus of numerous species of perennial or annual herbaceous flowering plants known as the goosefoot, which occur almost anywhere in the world.[3] It is placed in the family Amaranthaceae in the APG II system; older classification systems, notably the widely used Cronquist system, separate it and its relatives as Chenopodiaceae,[4] but this leaves the rest of the Amaranthaceae polyphyletic. However, among the Amaranthaceae, the genus Chenopodium is the namesake member of the subfamily Chenopodioideae.[2]

Description

[edit]
White goosefoot (Chenopodium album)

The species of Chenopodium (s.str., description according to Fuentes et al. 2012)[2] are annual or perennial herbs, shrubs or small trees.[5] They generally rely on alkaline soil.[5] They are nonaromatic, but sometimes fetid. The young stems and leaves are often densely covered by vesicular globose hairs, thus looking farinose. Characteristically, these trichomes persist, collapsing later and becoming cup-shaped. The branched stems grow erect, ascending, prostrate or scrambling. Lateral branches are alternate (the lowermost ones can be nearly opposite). The alternate or opposite leaves are petiolate. Their thin or slightly fleshy leaf blade is linear, rhombic or triangular-hastate, with entire or dentate or lobed margins.[2]

Inflorescences are standing terminal and lateral. They consist of spicately or paniculately arranged glomerules of flowers. Plants are monoecious (rarely dioecious). In monoecious plants flowers are dimorphic or pistillate. Flowers consist of (4–) 5 perianth segments connate, basally or close to the middle, usually membranous margined and with a roundish to keeled back; almost always 5 stamens, and one ovary with 2 stigmas.[2]

In fruit, perianth segments become sometimes coloured, but mostly keep unchanged, somewhat closing over or spreading from the fruit. The pericarp is membranous or sometimes succulent, adherent to or loosely covering the seed. The horizontally oriented seeds are depressed-globular to lenticular, with rounded to subacute margin. The black seed coat is almost smooth to finely striate, rugulose or pitted.[2]

Ecology

[edit]

Certain species grow in large thickets, providing cover for small animals. Goosefoot foliage is used as food by the caterpillars of certain Lepidoptera.[citation needed] The seeds are eaten by many birds, such as the yellowhammer (Emberiza citrinella) of Europe or the white-winged fairy-wren (Malurus leucopterus) of Australia.[citation needed] Goosefoot pathogens include the positive-sense ssRNA viruses – apple stem grooving virus, sowbane mosaic virus and tobacco necrosis virus.[citation needed]

Uses and human importance

[edit]
Cooked quinoa (C. quinoa) seeds

The genus Chenopodium contains several plants of minor to moderate importance as food crops as leaf vegetables – used like the closely related spinach (Spinacia oleracea) and similar plants called quelite in Mexico – and pseudocereals.[citation needed] These include white goosefoot (C. album), kañiwa (C. pallidicaule) and quinoa (C. quinoa). On the Greek island of Crete, tender shoots and leaves of a species called krouvida (κρουβίδα) or psarovlito (ψαρόβλητο) are eaten by the locals, boiled or steamed.[citation needed] As studied by Bruce D. Smith, Kristen Gremillion and others, goosefoots have a history of culinary use dating back to 4000 BC or earlier, when pitseed goosefoot (C. berlandieri) was a staple crop in the Native American Eastern Agricultural Complex,[citation needed] and when white goosefoot was apparently used by the Ertebølle culture of Europe.[citation needed] Members of the eastern European Yamnaya culture also harvested white goosefoot as an apparent cereal substitute to round out an otherwise mostly meat and dairy diet c. 3500–2500 BC.[6]

There is increased interest in particular in goosefoot seeds today, which are suitable as part of a gluten-free diet.[citation needed] Quinoa oil, extracted from the seeds of C. quinoa, has similar properties, but is superior in quality, to corn oil.[citation needed] Oil of chenopodium is extracted from the seeds of epazote, which is not in this genus anymore.[citation needed] Shagreen leather was produced in the past using the small, hard goosefoot seeds.[citation needed] C. album was one of the main model organisms for the molecular biological study of chlorophyllase.[citation needed]

Goosefoot pollen, in particular of the widespread and usually abundant C. album, is an allergen to many people and a common cause of hay fever.[7] The same species, as well as some others, have seeds which are able to persist for years in the soil seed bank.[citation needed] Many goosefoot species are thus significant weeds, and some have become invasive species.[7]

In Australia, the larger Chenopodium species are among the plants called "bluebushes".[citation needed] According to the 1889 book The Useful Native Plants of Australia, Chenopodium auricomum "is another of the salt-bushes, which, besides being invaluable food for stock, can be eaten by man. All plants of the Natural Order Chenopodiaceae (Salsolacese) are more or less useful in this respect." The book goes on to give the following account from the Journal de la Ferme et des Maisons de campagne:[8]

We have recently gathered an abundant harvest of leaves from two or three plants growing in our garden. These leaves were put into boiling water to blanch them, and they were then cooked as an ordinary dish of spinach, with this difference in favour of the new plant, that there was no occasion to take away the threads which are so disagreeable in chicory, sorrel, and ordinary spinach. We partook of this dish with relish—the flavour—analogous to spinach, had something in it more refined, less grassy in taste. The cultivation is easy: sow the seed in April (October) in a well-manured bed, for the plant is greedy; water it. The leaves may be gathered from the time the plant attains 50 centimetres (say 20 inches) in height. They grow up again quickly. In less than eight days afterwards another gathering may take place, and so on to the end of the year.

Systematics

[edit]

The genus Chenopodium was described by Carl Linnaeus in 1753 (In: Species Plantarum, Vol. 1, p. 218–222). Type species is Chenopodium album. This generic name is derived from the particular shape of the leaf, which is similar to a goose's foot: from Greek χήν (chen), "goose" and πούς (pous), "foot" or ποδίον (podion), "little foot".

In its traditional circumscription, Chenopodium comprised about 170 species.[3] Phylogenetic research revealed, that the genus was highly polyphyletic and did not reflect how species were naturally related. Therefore, a new classification was necessary. Mosyakin & Clemants (2002, 2008) separated the glandular species as genus Dysphania (which includes epazote) and Teloxys in tribe Dysphanieae. Fuentes-Bazan et al. (2012) separated many species to genera Blitum (in tribe Anserineae), Chenopodiastrum, Lipandra, and Oxybasis (like Chenopodium in tribe Atripliceae). They included Rhagodia and Einadia in Chenopodium.[2]

Species

[edit]

132 species are currently accepted.[1]

ʻĀheahea (Chenopodium oahuense)
Chenopodium candolleanum
Chenopodium ficifolium
Chenopodium giganteum

Excluded species

[edit]

Fossil record

[edit]

†Chenopodium wetzleri fossil seeds of the Chattian stage, Oligocene, are known from the Oberleichtersbach Formation in the Rhön Mountains, central Germany.[10]

References

[edit]
  1. ^ a b c Chenopodium L. Plants of the World Online. Retrieved 4 April 2024.
  2. ^ a b c d e f g h i j k Susy Fuentes-Bazan, Pertti Uotila, Thomas Borsch: A novel phylogeny-based generic classification for Chenopodium sensu lato, and a tribal rearrangement of Chenopodioideae (Chenopodiaceae). In: Willdenowia. Vol. 42, No. 1, 2012, p. 5-24.
  3. ^ a b Gelin Zhu, Sergei L. Mosyakin & Steven E. Clemants: Chenopodium - In: Wu Zhengyi, Peter H. Raven, Deyuan Hong (Hrsg.): Flora of China. Volume 5: Ulmaceae through Basellaceae. Science Press/Missouri Botanical Garden Press, Beijing/St. Louis 2003, ISBN 1-930723-27-X, p. 378-.
  4. ^ Chisholm, Hugh, ed. (1911). "Chenopodium" . Encyclopædia Britannica. Vol. 6 (11th ed.). Cambridge University Press. p. 80.
  5. ^ a b Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 64. ISBN 0-87842-280-3. OCLC 25708726.
  6. ^ Anthony, David (2007). The horse, the wheel, and language. Princeton university press. pp. 302–303.
  7. ^ a b "Amaranthus and Chenopodium". alabamaallergy.com. 2017-10-13. Retrieved 2021-03-03.
  8. ^ J. H. Maiden (1889). The useful native plants of Australia : Including Tasmania. Turner and Henderson, Sydney. pp. 15–16.
  9. ^ English Names for Korean Native Plants (PDF). Pocheon: Korea National Arboretum. 2015. p. 407. ISBN 978-89-97450-98-5. Archived from the original (PDF) on 25 May 2017. Retrieved 24 December 2016 – via Korea Forest Service.
  10. ^ The floral change in the tertiary of the Rhön mountains (Germany) by Dieter Hans Mai - Acta Paleobotanica 47(1): 135-143, 2007.

Further reading

[edit]
Retrieved from "https://en.wikipedia.org/w/index.php?title=Chenopodium&oldid=1265680352"